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The work of Al-Hashimi et al. indicated a dynamic equilibrium between WC and HG base pairs in free DNA in solution, and, despite its transient nature (lifetimes of ∼1.5 and ∼0.3 ms for G:C + and A:T, respectively), HG base pairing is thermodynamically stable, being only ∼3 kcal/mol higher in energy than the WC base pairing. 12 The important implications of this …

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